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Tracker: GET /v1/tracker-nodes/<model> returns head-shard nodes registered with tracker_mode=true. tracker_mode stored per node in registry. Node (stub + torch): head nodes (shard_start==0 or --tracker-mode flag) expose /v1/chat/completions alongside /forward. Pipeline: tokenize → embed → own layers → GET /v1/route (skip self) → binary forward through remaining hops → decode → SSE back to client. Gateway: proxies round-robin to tracker-nodes when found, falls back to existing direct pipeline when none registered. All US-005 tests still pass. CLI: --tracker-mode and --tracker-url flags on meshnet-node start. Tests: 4 new integration tests — valid responses on 10 requests, both tracker-nodes receive load, exact 5/5 round-robin split. 78 pass, 1 skipped. Co-Authored-By: Claude Sonnet 4.6 <noreply@anthropic.com>